1. Family: Lamiaceae Martinov
    1. Genus: Vitex L.
      1. Vitex fischeri Gürke

        This species is accepted, and its native range is South Sudan to S. Tropical Africa.

    [FZ]

    Flora Zambesiaca. Vol 8, Pt 7. Avicenniaceae, R. Fernandes. Nesogenaceae, M.A. Diniz. Verbenaceae, R. Fernandes. Lamiaceae, R. Fernandes. 2005.

    Type
    Type from Tanzania. Type from Angola.
    Habit
    Tree up to 15 m tall or more; bark shallowly fissured, ± smooth, greyish to pale yellow-brown; older branches yellow-brown; young branches and petioles orange-tawny tomentose Tree up to 15 m tall or more; bark shallowly fissured, ± smooth, greyish to pale yellow-brown; older branches yellow-brown; young branches and petioles orange-tawny tomentose.
    Leaves
    Leaves (4)5–7-foliolate; petioles (3)5–20 cm long, terete; median petiolules 0.5–2 cm long Leaves (4)5–7-foliolate; petioles (3)5–20 cm long, terete; median petiolules 0.5–2 cm long; leaflets 8–19 × 3–9 cm, median leaflet largest, elliptic, ovate to obovate, rounded to long acuminate at the apex, cuneate to rounded at the base, ± entire, somewhat scabrid-puberulous above to glabrescent, velvety-tomentose and sparsely to densely gland-dotted beneath and paler than the upper surface, nerves raised on both surfaces.
    Leaflets
    leaflets 8–19 × 3–9 cm, median leaflet largest, elliptic, ovate to obovate, rounded to long acuminate at the apex, cuneate to rounded at the base, ± entire, somewhat scabrid-puberulous above to glabrescent, velvety-tomentose and sparsely to densely gland-dotted beneath and paler than the upper surface, nerves raised on both surfaces
    Inflorescences
    Inflorescences axillary, of ± sturdy densely-flowered compound dichasia 5–16 cm long, pale yellow-tomentose; bracteoles 0.4–1 cm long, linear Inflorescences axillary, of ± sturdy densely-flowered compound dichasia 5–16 cm long, pale yellow-tomentose; bracteoles 0.4–1 cm long, linear.
    Calyx
    Calyx obconical, 5-lobed, lanate, sparsely gland-dotted, accrescent in fruit; tube 2–2.3 mm long; lobes 0.5–1.0 mm long, erect Calyx obconical, 5-lobed, lanate, sparsely gland-dotted, accrescent in fruit; tube 2–2.3 mm long; lobes 0.5–1.0 mm long, erect.
    Corolla
    Corollas 8–10 mm long, not gland-dotted, lanate or pubescent; tubes white, 5–6 mm long, curved; lobes mauve, at an obtuse angle to tube, middle lower lobe 3–5 × 2–3 mm Corollas 8–10 mm long, not gland-dotted, lanate or pubescent; tubes white, 5–6 mm long, curved; lobes mauve, at an obtuse angle to tube, middle lower lobe 3–5 × 2–3 mm.
    Stamens
    Stamens scarcely exserted from the corolla tube; filaments glabrous Stamens scarcely exserted from the corolla tube; filaments glabrous.
    Ovary
    Ovary obovoid, gland-dotted and pubescent toward the apex; apex truncate; style c. 6 mm long
    Fruits
    Drupe c. 1.3 × 0.7 cm, ellipsoid-obovoid, rounded at apex, the lower half enclosed by the enlarged calyx, purple-green to black with green spots at maturity; fruiting calyx cup-shaped , pubescent, not gland-dotted. Drupe c. 1.3 × 0.7 cm, ellipsoid-obovoid, rounded at apex, the lower half enclosed by the enlarged calyx, purple-green to black with green spots at maturity; fruiting calyx cup-shaped, pubescent, not gland-dotted.
    Ecology
    Miombo woodland, woodlands with Entandrophragma , Erythrophleum , Newtonia and Pachystela , wooded grassland and thickets, also on termite mounds; to 1220 m.
    Note
      V. fischeri is similar to V. madiensis subsp. milanjiensis in its inflorescences (densely flowered and shorter than the peduncles) in its small flowers and in leaflet shape and size.  However, it may be distinguished by stem colour (yellow-brown in V. fischeri , but dark purplish-brown in V. madiensis subsp. milanjiensis ) and by the leaflet lower surface indumentum (usually of dense curly hairs in V. fischeri and rather straight scattered hairs in V. madiensis ).   V. fischeri is distinguished from V. ferruginea , with which it shares the characters of petiolulate, long acuminate leaflets and leaves sometimes 7-foliolate, by the flower size (much larger in V. ferruginea ) and by the   leaflet lower surface indumentum (scattered small hairs in V. ferruginea ).
    Distribution
    District code: MOZ B, MOZ N, MOZ W. Zambia Also in Sudan, Uganda, Kenya, Tanzania, Dem. Rep. Congo and Angola.
    Pistil
    Ovary obovoid, gland-dotted and pubescent toward the apex; apex truncate; style c. 6 mm long.
    [FTEA]

    Verbenaceae, B. Verdcourt. Flora of Tropical East Africa. 1992

    Habit
    Savannah shrub or small to fairly large tree (1.8–)4.5–9 or sometimes to 15(–18) m. tall with rounded crown; bark grey to dark brown, shallowly striate and fissured, slash white to light brown with white sapwood; young branches and petioles densely clothed with velvety orange-brown hairs or yellow-tomentose.
    Leaves
    Leaves scented, 5-foliolate; leaflets oblong, ovate-elliptic to elliptic or slightly obovate-elliptic, 5–19 cm. long, 3–10 cm. wide, shortly to long-acuminate at the apex, the tip acute, cuneate to rounded at the base, entire, somewhat scabrid-puberulous above but later ± glabrous, velvety woolly tomentose beneath and with sparse to dense glands, the surface usually totally obscured but sometimes indumentum much sparser with glands and venation evident; lateral nerves rather numerous, up to ± 15 pairs; petioles 6.5–16.5 cm. long; petiolules 1–2(3–8) cm. long.
    Inflorescences
    Cymes axillary, extensive, many-flowered, up to 11(–15) cm. wide; peduncle 6–14 cm. long; pedicels 0–1 mm. long or that of central flower of cyme up to 12 mm.; all axes woolly-tomentose; bracts and bracteoles narrow, 0.3–2.2 cm. long, dark and ± glabrous inside, velvety outside.
    Calyx
    Calyx often tinged red, 3–4 mm. long, truncate or shallowly toothed, the teeth up to 1 mm. long.
    Corolla
    Corolla white with blue or mauve upper lip and lower cream or both blue; throat yellow; tube ± 3 mm. long, the lip 3.5 mm. long and lower lobes 2.5 mm. long and wide.
    Fruits
    Fruit black or purple-green to black with green spots, oblong-globose, 1–1.3 mm. long, 0.8–1 cm. wide, edible; calycine cup ± 1 cm. wide.
    Habitat
    Wooded grassland and thicket particularly on granite rocks, also rich bushland on termite mounds and Bridelia, Maesopsis, Albizia forest; 980–2080 m.
    Distribution
    K3 K5 K6 T1 T4 U1 U3 U4

    Images

    Distribution

    Native to:

    Angola, Kenya, Mozambique, Sudan, Tanzania, Uganda, Zambia, Zaïre

    Vitex fischeri Gürke appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Apr 3, 2014 Welwitsch, F.M.J. [5696], Angola K001067045 Unknown type material
    Dawkins, H.C. [415], Uganda 18107.000
    Grant, D. [846], Kenya K000192780
    Grant, D. [846], Kenya K000192781
    Grant, D. [846], Kenya K000192779

    First published in Bot. Jahrb. Syst. 18: 171 (1893)

    Accepted by

    • Meerts, P. (2018). Synopsis of the genus Vites (Lamiaceae) in the Democratic Republic of the Congo Plant Ecology and Evolution 151: 380-392.
    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Fischer, E., Rembold, K., Althof, A. & Obholzer, J. (2010). Annotated checklist of the vascular plants of Kakamega forest, Western province, Kenya Journal of East African Natural History 99: 129-226.
    • Figueiredo, E. & Smith, G.F. (2008). Plants of Angola Strelitzia 22: 1-279. National Botanical Institute, Pretoria.
    • Fernandes, R. & Diniz, M.A. (2005). Avicenniaceae, Nesogenaceae, Verbenaceae and Lamiaceae (subfams, Viticoideae and Ajugoideae) Flora Zambesiaca 8(7): 1-161. Royal Botanic Gardens, Kew.
    • Govaerts, R. (2003). World Checklist of Selected Plant Families Database in ACCESS: 1-216203. The Board of Trustees of the Royal Botanic Gardens, Kew.

    Literature

    Flora Zambesiaca
    • Kew Bull. 56 : 195 (2001).
    • F.T.E.A., Verbenaceae: 59 (1992).
    • White, For. Fl. N. Rhod.: 371 (1962).
    • Phytologia 5 : 335 (1955).
    • Brenan, Check-list For. Trees Shrubs Tang. Terr.: 642 (1949).
    • Bot. Jahrb. Syst. 62 , Beibl. 141: 60 (1928).
    • F.T.A. 5 : 330 (1900).
    Kew Backbone Distributions
    • Calane da Silva, M., Izdine, S. & Amuse, A.B. (2004). A Preliminary Checklist of the Vascular Plants of Mozambique: 1-184. SABONET, Pretoria.
    Flora of Tropical East Africa
    • F. White, Forest Flora of Northern Rhodesia p. 371 (1962).
    • Dale & Greenway, Kenya Trees and Shrubs p. 595, fig. 109/a,c,d (1961).
    • F. W. Andr., The Flowering Plants of the Sudan 3: 200 (1956).
    • W.J. Eggeling, Indigenous Trees of the Uganda Protectorate, ed. 2: 445 (1952).
    • J.P.M. Brenan, Check-lists of the Forest Trees and Shrubs of the British Empire no. 5, part II, Tanganyika Territory p. 643 (1949).
    • Pieper in A. Engler, Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 62, Beibl. 141: 60 (1928).
    • Bak. in Flora of Tropical Africa 5: 330 (1900).
    • Gürke in Die Pflanzenwelt Ost-Afrikas und der Nachbargebiete, Theile C: 339 (1895).
    • Gürke in A. Engler, Botanische Jahrbücher für Systematik, Pflanzengeschichte und Pflanzengeographie 18: 171 (1893)

    Sources

    Flora Zambesiaca
    Flora Zambesiaca
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0