1. Family: Poaceae Barnhart
    1. Genus: Zoysia Willd.
      1. Species: Zoysia matrella (L.) Merr.
        1. Zoysia matrella var. pacifica Goudsw.

        This variety is accepted, and its native range is Indo-China to Nansei-shoto and NW. Pacific.

    [GB]
    Habit
    Perennial; mat forming; clumped densely. Stolons present. Culms prostrate; 5-10 cm long; wiry. Lateral branches ample. Leaf-sheaths glabrous on surface. Leaf-sheath oral hairs ciliate; 2-5 mm long. Ligule a ciliate membrane. Leaf-blades filiform; 4-6 cm long; 1 mm wide.
    Inflorescences
    Inflorescence composed of racemes. Racemes 1; single; multilateral; 1-1.5 cm long; bearing 6-12 fertile spikelets on each. Spikelet packing lax. Spikelets ascending; solitary. Fertile spikelets pedicelled. Pedicels 1-1.6 mm long; scabrous; tip widened.
    Spikelets
    Spikelets comprising 1 fertile florets; without rhachilla extension. Spikelets lanceolate; laterally compressed; 2-3 mm long; 0.6 mm wide; falling entire.
    Fertile
    Spikelets comprising 1 fertile florets; without rhachilla extension. Spikelets lanceolate; laterally compressed; 2-3 mm long; 0.6 mm wide; falling entire.
    Glume
    Glumes one the lower absent or obscure; exceeding apex of florets; firmer than fertile lemma; shiny. Upper glume lanceolate; 2-3 mm long; 1 length of spikelet; coriaceous; without keels; 5 -veined. Upper glume primary vein obscure. Upper glume apex acute; muticous, or mucronate.
    Florets
    Fertile lemma oblong; 2-2.5 mm long; hyaline; without keel; 1 -veined. Palea absent or minute.
    Flowers
    Anthers 3; 0.6-0.8 mm long.
    Distribution
    Asia-temperate: eastern Asia. Asia-tropical: Indo-China, Malesia, and Papuasia. Pacific: south-central and northwestern.
    Reference
    Cynodonteae. Fl China 2009.

    Distribution

    Native to:

    Kazan-retto, Lesser Sunda Is., Maluku, Marianas, Myanmar, Nansei-shoto, Ogasawara-shoto, Philippines, Solomon Is., Taiwan, Thailand, Vietnam

    Introduced into:

    Bahamas, Benin, Burkina, Cayman Is., Dominican Republic, Ecuador, El Salvador, Florida, Gabon, Galápagos, Haiti, Hawaii, Honduras, India, Ivory Coast, Mexico Northeast, Nicaragua, Puerto Rico, West Himalaya

    Zoysia matrella var. pacifica Goudsw. appears in other Kew resources:

    First published in Blumea 26: 172 (1980)

    Accepted by

    • Jaramillo Díaz, P. & Guézou, A. (2017). CDF Checklist of Galapagos Vascular Plants - FCD Lista de especies de Plantas Vasculares de Galápagos http://www.darwinfoundation.org/datazone/checklists/vascular-plants/.
    • Valdés-Reyna, J. Villaseñor, J.L., Encina-Domínguez, J.A. & Oritz, E. (2015). The grass family (Poaceae) in Coahuila, Mexico: diversity and distribution Botanical Sciences 93: 119-129.
    • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Clayton, W.D., Harman, K.T. & Williamson, H. (2006). World Grass Species - Synonymy database The Board of Trustees of the Royal Botanic Gardens, Kew.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
    • Turner, I.M., Xing, F. & Corlett, R.T. (2000). An annotated check-list of the vascular plants of the South China Sea and its shores The Raffles bulletin of Zoology, Suppl. 8: 23-116.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.

    Literature

    Kew Backbone Distributions
    • Hall, D.W. (2019). Grasses of Florida: 1-353. University press of Florida.
    • Jaramillo Díaz, P. & Guézou, A. (2017). CDF Checklist of Galapagos Vascular Plants - FCD Lista de especies de Plantas Vasculares de Galápagos http://www.darwinfoundation.org/datazone/checklists/vascular-plants/.
    • Valdés-Reyna, J. Villaseñor, J.L., Encina-Domínguez, J.A. & Oritz, E. (2015). The grass family (Poaceae) in Coahuila, Mexico: diversity and distribution Botanical Sciences 93: 119-129.
    • Girmansyah, D. & al. (eds.) (2013). Flora of Bali an annotated checklist: 1-158. Herbarium Bogorensis, Indonesia.
    • Thiombiano, A., Schmidt, M., Dressler, S., Ouédraogo, A., Hahn, K. & Zizka, G. (2012). Catalogue des plantes vasculaires du Burkina Faso Boissiera 65: 1-391.
    • Acevedo-Rodríguez, P. & Strong, M.T. (2012). Catalogue of seed plants of the West Indies Smithsonian Contributions to Botany 98: 1-1192.
    • Kandwal, M.K. & Gupta, B.K. (2009). An update on grass flora of Uttarkhand Indian Journal of Forestry 32: 657-668.
    • Nelson Sutherland, C.H. (2008). Catálogo de las plantes vasculares de Honduras. Espermatofitas: 1-1576. SERNA/Guaymuras, Tegucigalpa, Honduras.
    • Wu, Z. & Raven, P.H. (eds.) (2006). Poaceae Flora of China 22: 1-733. Missouri Botanical Garden Press, St. Louis.
    • Sosef, M.S.M. & al. (2006). Check-list des plantes vasculaires du Gabon Scripta Botanica Belgica 35: 1-438.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Kress, W.J., DeFilipps, R.A., Farr, E. & Kyi, D.Y.Y. (2003). A Checklist of the Trees, Shrubs, Herbs and Climbers of Myanmar Contributions from the United States National Herbarium 45: 1-590.
    • Flora of North America Editorial Committee (2003). Flora of North America North of Mexico 25: 1-781. Oxford University Press, New York, Oxford.
    • Aké Assi, L. (2002). Flore de la Côte-d'Ivoire: catalogue systématique, biogéographie et écologie. II Boissiera 58: 1-401.
    • H?, P.-H. (2000). Câyc? Vi?tnam. An Illustrated flora of Vietnam, ed. 2, 3: 1-1020. Pham-hoang Ho, Montréal.
    • Jørgensen, P.M. & León-Yánez, S. (eds.) (1999). Catalogue of the Vascular Plants of Ecuador Monographs in Systematic Botany from the Missouri Botanical Garden 75: i-viii, 1-1181.
    • Davidse, G. & al. (eds.) (1994). Flora Mesoamericana 6: 1-543. Universidad Nacional Autónoma de México, México, D.F.
    • Karthikeyan, S., Jain, S.K., Nayar, M.P. & Sanjappa, M. (1989). Florae Indicae Enumeratio: Monocotyledonae: 1-435. Botanical Survey of India, Calcutta.
    • Kobayashi, S. & Ono, M. (1987). A Revised List of Vascular Plants Indigenous and Introduced to the Bonin (Ogasawara) and the Volcano (Kazan) Islands Ogasawara Research 13: 1-55.

    Sources

    GrassBase - The Online World Grass Flora
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2020. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0