1. Family: Cannabaceae Martinov
    1. Genus: Celtis L.
      1. Celtis toka (Forssk.) Hepper & J.R.I.Wood

        This species is accepted, and its native range is W. Tropical Africa to Arabian Peninsula.

    [FWTA]

    Ulmaceae, Hutchinson and Dalziel. Flora of West Tropical Africa 1:2. 1958

    Habit
    A large spreading tree, to 80 ft. high and 15 ft. girth
    Ecology
    Mainly in the drier savannah regions.
    [FTEA]

    Ulmaceae, R. M. Polhill. Flora of Tropical East Africa. 1966

    Habit
    Large spreading deciduous monoecious tree, up to 25 m. high, with tall narrow buttresses; bark light grey, smooth or scaly.
    Twigs
    Young twigs pubescent.
    Leaves
    Leaf-blades broadly ovate, 4.6–7.8 cm. long, 3.5–4.8 cm. wide, shortly acuminate, rounded to subcordate at the base, unequal-sided, usually entire, sometimes toothed in the upper half, scabrous, subglabrous with a few hairs on the veins beneath and in the vein-angles, 3–5-nerved from the base, with the basal lateral nerves ascending well into the upper half and the upper prominent lateral nerves 1–3 on each side of the midrib; petiole 4–6 mm. long.
    Stipules
    Stipules linear, 4–5 mm. long, pubescent.
    Inflorescences
    Cymes 1–2 cm. long in flower, up to 4 cm. in fruit, axillary or at the nodes below, with many clustered ♂ flowers and, the upper ones in particular, with several terminal ♀, hermaphroditeflowers on ± 3 mm. long pedicels.
    Calyx
    Sepals 5, 2–2.5 mm. long, pubescent.
    Pistil
    Ovary shortly pubescent with a ring of longer hairs at the base; styles. 2–3-branched, 3–5 mm. long.
    Fruits
    Fruits subglobose, 8–11 mm. long, 7.5–10 mm. across, brownish, puberulous or scabrous.
    Habitat
    Dry riverine forest and deciduous woodland; 600–1200 m.
    Distribution
    U1 Sudan Republic, west to Senegal and east to Arabia

    Images

    Distribution

    Native to:

    Benin, Burkina, Cameroon, Central African Repu, Chad, Eritrea, Ethiopia, Gambia, Ghana, Guinea, Ivory Coast, Mali, Mauritania, Niger, Nigeria, Senegal, Somalia, Sudan, Togo, Uganda, Yemen

    Celtis toka (Forssk.) Hepper & J.R.I.Wood appears in other Kew resources:

    Date Reference Identified As Barcode Type Status
    Chevalier, A. [1136], Mali Celtis integrifolia K000386754
    Starin, E.D. [1] Celtis integrifolia 40496.000
    Headley, P.M. [228], Ethiopia Celtis integrifolia 6049.060
    Roger [s.n.], Senegal Celtis integrifolia K000243028
    Dubois, R. [204], Mali Celtis integrifolia K000386753

    First published in Kew Bull. 38: 86 (1983)

    Accepted by

    • Darbyshire, I., Kordofani, M., Farag, I., Candiga, R. & Pickering, H. (eds.) (2015). The Plants of Sudan and South Sudan: 1-400. Kew publishing, Royal Botanic Gardens, Kew.
    • Kalema, J. & Beentje, H. (2012). Conservation checklist of the trees of Uganda: 1-235. Kew Publishing, Royal Botanic Gardens, Kew.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Govaerts, R. (1999). World Checklist of Seed Plants 3(1, 2a & 2b): 1-1532. MIM, Deurne.
    • Thulin, M. (ed.) in Thulin, M. (ed.) (1999). Flora of Somalia 2: 1-303. The Royal Botanic Gardens, Kew.
    • Wood, J.R.I. (1997). A handbook of the Yemen Flora: 1-434. Royal Botanic Gardens, Kew.
    • Hedberg, I. & Edwards, S. (eds.) (1989 publ. 1990). Flora of Ethiopia and Eritrea 3: 1-659. The National Herbarium, Addis Ababa University, Ethiopia & The Department of Systematic Botany, Upps.

    Literature

    Kew Backbone Distributions
    • Onana, J.M. (2011). The vascular plants of Cameroon a taxonomic checklist with IUCN assessments: 1-195. National herbarium of Cameroon, Yaoundé.
    • Akoègninou, A., van der Burg, W.J. & van der Maesen, L.J.G. (eds.) (2006). Flore Analytique du Bénin: 1-1034. Backhuys Publishers.
    • Thulin, M. (ed.) in Thulin, M. (ed.) (1999). Flora of Somalia 2: 1-303. The Royal Botanic Gardens, Kew.
    • Lebrun, J.p., Toutain, B., Gaston, A. & Boudet, G. (1991). Catalogue des Plantes Vasculaires du Burkina Faso: 1-341. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Boulvert, Y. (1977). Catalogue de la Flore de Centrafrique 3: 1-89. ORSTROM, Bangui.
    • Peyre de Fabregues, B. & Lebrun, J.-P. (1976). Catalogue des Plantes Vascularies du Niger: 1-433. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.
    • Lebrun, J.-P., Audru, J., Gaston, A. & Mosnier, M. (1972). Catalogue des Plantes Vasculaires du Tchad Méridional: 1-289. Institut d' Elevage et de Médecine Vétérinaire des Pays Tropicaux, Maisons Alfort.

    Sources

    Flora of Tropical East Africa
    Flora of Tropical East Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Flora of West Tropical Africa
    Flora of West Tropical Africa
    http://creativecommons.org/licenses/by-nc-sa/3.0

    Herbarium Catalogue Specimens
    'The Herbarium Catalogue, Royal Botanic Gardens, Kew. Published on the Internet http://www.kew.org/herbcat [accessed on Day Month Year]'. Please enter the date on which you consulted the system.
    Digital Image © Board of Trustees, RBG Kew http://creativecommons.org/licenses/by/3.0/

    Kew Backbone Distributions
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0

    Kew Names and Taxonomic Backbone
    The International Plant Names Index and World Checklist of Selected Plant Families 2019. Published on the Internet at http://www.ipni.org and http://apps.kew.org/wcsp/
    © Copyright 2017 International Plant Names Index and World Checklist of Selected Plant Families. http://creativecommons.org/licenses/by/3.0