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This species is accepted, and its native range is Europe to Xinjiang and Himalaya, Macaronesia to N. Africa.
Lolium arundinaceum


Bernal, R., G. Galeano, A. Rodríguez, H. Sarmiento y M. Gutiérrez. 2017. Nombres Comunes de las Plantas de Colombia.

cañuela, paja de páramo

Morphology General Habit
Perennial; caespitose; clumped moderately. Cataphylls inconspicuous. Rhizomes absent, or short; fleshy; not obviously scaly. Basal innovations extravaginal. Culms robust; straight, or curved; 45-180 cm long; without nodal roots. Culm-internodes terete. Culm-nodes without exudate; glabrous. Lateral branches lacking. Leaves basal and cauline; 3-5 per branch. Leaf-sheaths tight; unthickened at base; open for most of their length; without keel; striately veined; glabrous on surface. Leaf-sheath oral hairs ciliate. Leaf-sheath auricles falcate. Ligule an eciliate membrane; 1-2 mm long; membranous, or scarious; white; erose. Leaf-blade base symmetrical. Leaf-blades straight, or flexuous; linear in section; 10-60 cm long; 3-12 mm wide; firm; mid-green. Leaf-blade midrib conspicuous. Leaf-blade venation distinct; with subepidermal sclerenchyma strands similar in size; with subepidermal sclerenchyma attached to veins above and below; without layer of subepidermal sclerenchyma masking vein striation. Leaf-blade surface glabrous. Leaf-blade margins smooth; glabrous. Leaf-blade apex attenuate.
Morphology Reproductive morphology Inflorescences
Inflorescence a panicle; with 1 peduncles per sheath. Peduncle straight, or flexuous; terete; eglandular; glabrous. Panicle open; lanceolate, or ovate; continuous; dense, or loose; straight; 10-50 cm long. Primary panicle branches ascending, or spreading; 2 -nate; moderately divided; bearing 3-8 fertile spikelets on each lower branch (on shorter branch); bearing spikelets almost to the base. Panicle branches straight, or arcuate; angular; scabrous. Spikelets solitary. Fertile spikelets pedicelled. Pedicels 3-8 mm long.
Morphology Reproductive morphology Inflorescences Spikelets
Spikelets comprising 6-8(-10) fertile florets; with diminished florets at the apex. Spikelets elliptic, or oblong; laterally compressed; 9-12(-15) mm long; breaking up at maturity; disarticulating below each fertile floret. Spikelet callus glabrous. Rhachilla internodes scaberulous. Floret callus glabrous.
Spikelets comprising 6-8(-10) fertile florets; with diminished florets at the apex. Spikelets elliptic, or oblong; laterally compressed; 9-12(-15) mm long; breaking up at maturity; disarticulating below each fertile floret. Spikelet callus glabrous. Rhachilla internodes scaberulous. Floret callus glabrous.
Morphology Reproductive morphology Inflorescences Bracts Glume
Glumes persistent; dissimilar; subequal in width; shorter than spikelet. Lower glume lanceolate; 3.3-5.5 mm long; 0.6 length of upper glume; herbaceous, or scarious; mid-green; without keels; 1 -veined. Lower glume lateral veins absent. Lower glume apex obtuse. Upper glume lanceolate, or oblong; 4.5-7 mm long; 0.7-0.8 length of adjacent fertile lemma; chartaceous; mid-green; without keels; 3 -veined. Upper glume apex acute.
Morphology Reproductive morphology Flowers Florets
Fertile lemma ovate; symmetrical; 6-9 mm long; chartaceous; mid-green; without keel; 5 -veined. Lemma surface scaberulous. Lemma apex acute; muticous, or awned; 1 -awned. Principal lemma awn stiff; 0-4 mm long overall. Palea 1 length of lemma; 2 -veined. Palea keels scaberulous. Apical sterile florets resembling fertile though underdeveloped.
Morphology Reproductive morphology Flowers
Anthers 3; 3-4 mm long. Ovary glabrous.
Morphology Reproductive morphology Fruits
Caryopsis with adherent pericarp; glabrous. Hilum linear; 0.5 length of caryopsis.
Europe: northern, central, southwestern, southeastern, and eastern. Africa: north, Macaronesia, northeast tropical, east tropical, and south. Asia-temperate: Siberia, Soviet Middle Asia, Caucasus, western Asia, Arabia, China, and eastern Asia. Asia-tropical: India and Papuasia. Australasia: Australia and New Zealand. Pacific: north-central. North America: Subarctic, western Canada, eastern Canada, northwest USA, north-central USA, northeast USA, southwest USA, and Mexico. South America: Mesoamericana, western South America, Brazil, and southern South America. Antarctic: Subantarctic islands.
Poeae. Stancik 2005.


Bernal, R., Gradstein, S.R. & Celis, M. (eds.). 2015. Catálogo de plantas y líquenes de Colombia. Instituto de Ciencias Naturales, Universidad Nacional de Colombia, Bogotá.

Naturalizada y adventicia en Colombia; Alt. 2500 - 3000 m.; Andes.
Morphology General Habit
No Evaluada


Gramineae, W. D. Clayton, S. M. Phillips & S. A. Renvoize. Flora of Tropical East Africa. 1974

Morphology General Habit
Tufted perennial; culms 45–180 cm. high.
Morphology Leaves
Leaf-blades 10–60 cm. long and 3–12 mm. wide, flat, with minutely hairy falcate auricles; leaf-sheaths glabrous.
Morphology Reproductive morphology Inflorescences
Panicle lanceolate to ovate, 10–50 cm. long, loose or contracted.
Morphology Reproductive morphology Inflorescences Spikelets
Spikelets closely 3–10-flowered, elliptic to oblong, 10–18 mm. long; lower glume 3–6 mm. long, the upper 4.5–7 mm. long; lemmas narrowly ovate, 6–9 mm. long, awnless or with a fine bristle up to 4 mm. long from just below the hyaline tip; palea as long as the lemma; ovary glabrous; anthers 3–4 mm. long.
Upland streamsides; 2300–3000 m.
K3 Europe and temperate Asiawidely introduced elsewhere

Doubtfully present in:


Native to:

Afghanistan, Albania, Algeria, Austria, Azores, Baleares, Baltic States, Belarus, Belgium, Bulgaria, Canary Is., Central European Rus, Cyprus, Czechoslovakia, Denmark, East Aegean Is., East European Russia, East Himalaya, Finland, France, Germany, Great Britain, Greece, Hungary, Iran, Iraq, Ireland, Italy, Kazakhstan, Kirgizstan, Kriti, Krym, Lebanon-Syria, Libya, Morocco, Netherlands, North Caucasus, Northwest European R, Norway, Pakistan, Palestine, Poland, Portugal, Romania, Sardegna, Sicilia, South European Russi, Spain, Sweden, Switzerland, Tadzhikistan, Transcaucasus, Tunisia, Turkey, Turkey-in-Europe, Turkmenistan, Ukraine, Uzbekistan, West Himalaya, West Siberia, Xinjiang, Yugoslavia

Introduced into:

Alabama, Alaska, Alberta, Argentina Northeast, Argentina South, Arizona, Arkansas, Bolivia, Brazil South, British Columbia, California, Cape Provinces, Chatham Is., Chile Central, Chile North, Chile South, China North-Central, China South-Central, China Southeast, Colombia, Colorado, Connecticut, Costa Rica, Delaware, District of Columbia, Dominican Republic, Ecuador, Egypt, Ethiopia, Florida, Free State, Georgia, Guatemala, Haiti, Hawaii, Idaho, Illinois, Inner Mongolia, Iowa, Jamaica, Japan, Juan Fernández Is., Kamchatka, Kansas, Kentucky, Kenya, Kerguelen, Korea, Kuril Is., KwaZulu-Natal, Louisiana, Madeira, Maine, Manchuria, Maryland, Massachusetts, Mexico Central, Mexico Northeast, Michigan, Minnesota, Mississippi, Missouri, Montana, Nebraska, Nevada, New Brunswick, New Guinea, New Hampshire, New Mexico, New South Wales, New York, New Zealand North, Newfoundland, North Carolina, Nova Scotia, Ohio, Oklahoma, Ontario, Oregon, Pennsylvania, Qinghai, Québec, Sakhalin, Saskatchewan, Saudi Arabia, South Australia, South Carolina, South Dakota, Taiwan, Tasmania, Tennessee, Texas, Tristan da Cunha, Uruguay, Utah, Vermont, Victoria, Virginia, Washington, West Virginia, Western Australia, Wisconsin, Wyoming, Yukon

Lolium arundinaceum (Schreb.) Darbysh. appears in other Kew resources:

Date Reference Identified As Barcode Type Status
Oct 1, 1976 Rechinger, K.H. [1144], Iran Festuca arundinacea K000913147 Unknown type material
Pitard, C. J. [1254], Morocco Festuca arundinacea K000345316
Pitard, C. J. [1255], Morocco Festuca arundinacea K000345317
Cope, T.A. [RBG 401], United Kingdom Festuca arundinacea K000914496

First published in Novon 3: 241 (1993)

Not accepted by

  • Clayton, W.D., Harman, K.T. & Williamson, H. (2006). World Grass Species - Synonymy database The Board of Trustees of the Royal Botanic Gardens, Kew. [Cited as Festuca arundinacea.]


Catálogo de Plantas y Líquenes de Colombia

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Kew Backbone Distributions

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